1) Those trees being very far away from what an E. vernicosa looks like (E. vernicosa is a dwarf shrub with a mallee habit).
2) Those trees having flower buds very alike to not just E. gunnii, but allied species of E. gunnii (namely, taxa like archerii, divaricata, gunnii, urnigera, etc.)
3) Those trees having flower buds looking different to those of E. subcrenulata, which in turn, makes them have flower buds looking different to those of E. johnstonii. Those two are the only ones who have a "large tree" outlook for the Yellow Gum cline.
Surely, bark is not alike any "classic E. gunnii" I have seen by now. So, the next question for me is "why?" If the answer was "because it is an hybrid" (something we do not know yet), then:
1) Both parent species would have flower buds and capsules in groups of 3. Otherwise, the trees at Villa Thuret should show intermediate features (more than 3 in some inflorescences and infrutescences)
2) The hybrid would very likely not be an E. gundal (E. gunnii x dalrympleana, the most common hybrid in France for E. gunnii) or flower buds would look different.
3) One of the parent trees would possibly be part of the Cider Gum Complex (E. gunnii and allied species).
4) One of the parent trees would have rough bark for more or less half the tree height, and must be compatible with the other parent tree. So, almost surely it would also belong to Symphyomyrtus, very probably Section Maidenaria if the other parent is part of the Cider Gum Complex ("Superspecies Gunnii"). This is because Inter-Sectional hybrids are very difficult. With less security than the previous, but possibly, it would not be part of Series Viminales (E. dalrympleana hybrid with E. gunnii has different buds) or Series Johnstonianae (E. subcrenulata has different buds) if the other parent is part of the Cider Gum Complex.
5) One of the parent species cannot yet be discarded to be an hybrid. (This is the trans-generational hybrid hypothesis you well mentioned). Something like: (Species A) x (Species B x Species C), without discarding A=B or A=C.
So, I think we think very alike George! And the good thing is the range of "probable species" involved in this taxonomic mine field is now quite less than 700!
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