It has occurred to me that a similar thing happened later on, when Charlotte married her first cousin, Prince Felix of Bourbon-Parma -- who, despite being a member of a long-deposed dynasty, was a Royal Highness as a male-line descendant of King Louis XIV of France (itself a long-abolished monarchy). Accordingly, their own male-line descendants have been elevated to that style.
Nevertheless, isn't the style of dubious validity, in light of the fact that it comes from the non-reigning royal house of Bourbon? Yet, it is being applied to members of a reigning grand duchy ...
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